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54 J.C. SPARKS, R.E. MASTERS and M.E. PAYTON <br />method can also be used to sample all stems <br />in an area regardless of size, but produced <br />less accurate estimates than those of the FRP- <br />AC or FRP-HA methods when samp-ling <br />small stems. However, the QUAD provided <br />more accurate estimates of larger stems than <br />did both FRP methods. The QUAD tended <br />to be less time -efficient than the FRP <br />methods and required the use of a transit <br />and tape measure for accurate plot layout <br />or a large sampling quadrat, which can be <br />cumbersome to transport and set up in <br />dense stands. The QUAD method requires <br />considerable care so that stems are not <br />counted twice. <br />Although the VRP method was one of <br />the fastest methods, its use should be limited <br />to primarily larger stems with a DBH >11.43 <br />cm. The VRP method produced poor and <br />wide ranging accuracies on smaller stems <br />(<11.43 cm), overestimating dense stands <br />and underestimating low -density stands. <br />The VRP also tended to underestimate <br />overall stem density of stems >11.43 cm, but <br />estimates were a consistent percent dif- <br />ference across all stand densities and may <br />be reduced if additional samples were taken. <br />The BT produced poor estimates on <br />small stems, but relatively accurate esti- <br />mates on larger stems. Although it can be <br />used to sample all stems regardless of size, <br />this method was not very time -efficient, <br />primarily because transects must be set up <br />and measured before sampling can begin. <br />This method would be best suited to long <br />transects, with the aid of a compass and a <br />device to measure distance traveled. The BT <br />method may be best applied in transition <br />zones or gradients where the vegetation <br />changes in composition and density (6). <br />The PCQ method was quite time - <br />efficient, but was applied only to larger <br />stems. The PCQ method tended to under- <br />estimate actual stem density in all stands, <br />but like the VRP, the PCQ method also <br />maintained a consistent percent difference <br />in observed versus estimated stem densities <br />across all stands regardless of overall stem <br />density. That distance techniques may give <br />biased estimates, depending on tree spatial <br />distributions, is an inherent disadvantage. <br />Shape: FRPs with a circular shape tended <br />to produce the most accurate results on <br />smaller stems (2.54 to 11.42 cm), especially <br />when applied in dense, evenly distributed <br />stands. This can be attributed to the small <br />perimeter to area ratio in a circle; therefore, <br />an observer is less likely to have to make a <br />decision to tally a stem or not because it <br />intersects the plot (17). Also, the rotating <br />radius of the plot allows an observer to <br />ensure all stems are tallied and are only <br />tallied once. When using a large quadrat <br />(10 m x 10 m) it is easy to miss stems or to <br />tally them twice (6,18). The FRP methods <br />or circular -plot sampling methods are quick <br />and simple to apply in areas with low to <br />moderate density of stems or areas with low <br />vegetation, but become awkward in dense, <br />shrubby communities (6,18). Smaller FRP <br />or square -plot methods can be used in <br />stands with dense vegetation (18). <br />Size: It is essential for quadrat size to be <br />adapted to the characteristics of the vege- <br />tation being sampled (18). The greater the <br />species diversity and the more hetero- <br />geneous life forms found in a community, <br />the larger the quadrat size needed to <br />adequately characterize the community (18). <br />Regardless of shape, perimeter to area ratios <br />decrease with an increase in quadrat size <br />(17). Based on the diversity found in many <br />forested communities and on perimeter to <br />area ratios, larger circular plots would be <br />most appropriate (17). <br />CONCLUSIONS <br />It is important for managers to know the <br />general characteristics of the stand they are <br />attempting to sample because no single <br />method is efficient for sampling all stem <br />sizes and densities. Sample procedures <br />provide only an estimate, and this estimate <br />may be smaller or greater than actual stem <br />density depending on the sample method <br />applied and stand density. Methods should <br />be chosen that reflect unbiased estimates of <br />density given various stand conditions and <br />distributions. <br />Circular and fixed area methods, such <br />as the FRP-AC and FRP-HA, were the most <br />Proc. Okla. Acad. Sci. 82:49-56(2002) <br />